" We knew that the population was expanding , both in numbers and spatially on the landscape , " said joshua schmidt , a biometrician for the national park service inventory and monitoring program and a co-author of the study .

“我们知道人口的膨胀无论在数量和空间景观”施密特约书亚说对于国家公园的服务biometrician库存和监控程序和本研究合著者。